ExpressionUUID

is an option for Cell and Notebook that holds the assigned unique UUID string.

Details

ExpressionUUID is managed automatically by the front end and cannot be manually assigned, queried or inherited as a typical notebook front end option.
All notebooks and cells have an ExpressionUUID option written to the file when the file is saved.
Functions that read the contents of open notebook windows such as NotebookGet and NotebookRead will not include the value of ExpressionUUID in their results. This helps to ensure that a UUID does not get reused when programmatically modifying notebooks using such commands.
The UUID designated by ExpressionUUID is the same as the one used in the first argument of NotebookObject and CellObject to uniquely identify a given instance of a cell.

Query the ExpressionUUID of a cell using Developer`CellInformation:

Use NotebookLocate to locate the cell with the given ExpressionUUID:

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